The Grünbach Formation is an Austrian geological formation that dates to the early Campanian age of the Late Cretaceous. it forms part of the Gosau Group, and represents a marine regression event, representing a coastal/brackish environment, being underlain by the marine carbonate Maiersdorf Formation and overlain by the deep marine siliciclastic Piesting Formation. The main lithology is clay, marl, siltstone and sandstone, with a minor conglomerate component. Coal seams have also been noted.[1][2] It is notable for its fossils including those of dinosaurs and plants.
Vertebrate paleofauna
All remains have been found at a single locality, which is a thin marl seam in the Konstantin mining tunnel, within the Felbering Mine in the Neue Welt area north west of Muthmannsdorf in Lower Austria.[3] The initial remains were discovered in 1859 after an ornithopod tooth was found in a piece of coal in a dump outside the mine by Professor Ferdinand Stoliczka, and the productive seam discovered thereafter. The first material was described by Emanuel Bunzel in 1871[4] and then additional material was described by Harry Seeley in 1881.[5] Due to mining activity in the area ceasing at the end of the 19th century, no additional remains have been recovered since.
Squamates
| Squamates of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| Araeosaurus
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A. gracilis
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Muthmannsdorf
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Vertebra[5]
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| Crocodyliformes of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| Doratodon
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D. carcharidens
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Muthmannsdorf
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"incomplete mandible, a fragmentary right maxilla, a parietal fragment, and isolated teeth"[6]
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| Eusuchia
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Indeterminate
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Muthmannsdorf
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"mandible fragment and some postcranial material"
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Choristoderes
| Choristoderes of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| Choristodera
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Indeterminate
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Muthmannsdorf
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"Two platycoelous vertebral centra"[7]
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Turtles
| Turtles of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| Dortokidae[8]
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Indeterminate
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Muthmannsdorf
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Shell fragments
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| cf. Kallokibotioninae
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Indeterminate
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Muthmannsdorf
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Shell fragments
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Pterosaurs
| Pterosaurs of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| "Ornithocheirus"
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"O". buenzeli
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Muthmannsdorf
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Remains found not diagnostic, remains referred to Azdarchidae and Pterodactyloidea indet.
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| Azdarchidae
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Indeterminate
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Muthmannsdorf
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"Proximal portion of a humerus"[9]
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Previously referred to "O". buenzeli
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| Pterodactyloidea
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Indeterminate
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Muthmannsdorf
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"Articular region of a lower jaw, crushed phalangeal fragments"
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Previously referred to "O". buenzeli
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Non-avian dinosaurs
| Dinosaurs of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic position
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Abundance
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Notes
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Images
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| Mochlodon
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M. suessi
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Muthmannsdorf
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Cranial and postcranial elements
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| Struthiosaurus
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S. austriacus
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Muthmannsdorf
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Cranial and postcranial elements
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| Tetanurae
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Indeterminate
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Muthmannsdorf
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"Two fragmentary teeth"
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Referred to by the dubious name ‘Megalosaurus pannoniensis’[5] Nearly identical to teeth known from the Csehbánya Formation[10]
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Flora
Most of these specimens were recovered from mining dumps near Grünbach am Schneeberg in lower Austria. The flora of the formation is considered to represent that of a high humidity subtropical climate, typical of the Euro-Sinian phytogeographical region.[11][12][13]
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Color key
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Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
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Bryophytes
| Bryophytes of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Stereisporites
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Indeterminate
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Spore, representative of Sphagnaceae
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Lycopodiophyta
| Lycopodiophytes of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Camarozonosporites
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Indeterminate
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Spore, representative of Lycopodiaceae
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| Retitriletes
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Indeterminate
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Spore, representative of Lycopodium
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Pteridophytes
| Pteridophytes of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Appendicisporites
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A. tricuspidatus
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Spore, probable affinities with Schizaeaceae
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| Cicatricosisporites
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Indeterminate
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Spore, probable affinities with Schizaeaceae
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| Cingulatisporites
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Indeterminate
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Spore, affinities with Anemia
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| Cladophlebis
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C. gosauensis
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| Coniopteris
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?
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| Cyathidites
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Indeterminate
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Spore
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| Echinatisporites
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Indeterminate
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Spore
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| Equisetites
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Indeterminate
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A horsetail
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| Gosauopteris
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G. danaeoides
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Probable affinities with Marattiaceae
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| Gleicheniidites
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G. senonicus
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Spore
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| Leiotriletes
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Indeterminate
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Spore, representative of Lygodium
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| Marsileaceaephyllum
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M. campanica
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Marsileaceae, very similar to extant Marsilea
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| Matonisporites
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Indeterminate
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Spore, Found in situ with Monheimia
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| Microphyllopteris
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M. austriaca
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| Microtaenia
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M. austriaca
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| Monheimia
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M. ungerii
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affinities with Matoniaceae, particularly Phanerosorus
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| Polypodiaceoisporites
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Indeterminate
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Spore
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| Raphaelia
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R. lobifolia
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| Sphenopteris
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S. gruenbachiana, S. ungeri
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Probable affinities with Schizaeaceae
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Gymnosperms
| Gymnosperms of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Geinitzia
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G. formosa, G. reichenbachii
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Pinopsida, Taxodioid affinity
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| cf. Lindleycladus
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L. lanceolatus
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| Nilsonia
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N. cf. holyi
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Bennettitales
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| Pagiophyllum
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Indeterminate
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| Pityosporites
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Indeterminate
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Pollen
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| Podozamites
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Indeterminate
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| Taxiodiaceaepollenites
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Indeterminate
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Pollen, at least 2 taxa
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Angiosperms
Monocots
| Monocots of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Arecipites
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Indeterminate
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|
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Pollen, representative of Palmae
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| Lysichiton
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L. austriacus
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|
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Araceae, genus extant
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| Gruenbachia
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G. pandanoides
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Pandanaceae, reproductive organs of Pandanites
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| Pandanites
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P. trinervis
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Pandanaceae
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| Sabalites
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S. longirhachis
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Palm
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| Spinizonocolpites
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Indeterminate
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Pollen, representative of extant Nypa palm
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| Theiaiphyllum
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T. kollmannii
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Dicots
| Dicots of the Grünbach Formation
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| Genus
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Species
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Location
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Stratigraphic Position
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Material
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Notes
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Images
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| Brasenites
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B. krasserii
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affinity with Brasenia
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| Celastrophyllum
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Indeterminate, C. johannae
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Hamamelidae
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| Ceratoxylon
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Indeterminate
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Reproductive structure
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| Clavatipollenites
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Indeterminate
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|
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Pollen, representative of Chloranthaceae
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| Complexiopollis
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Indeterminate
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Pollen, affinities with Myricaceae
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| Compositiphyllum
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C. serratum
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| Debeya
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D. insignis
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| Dicotylophyllum
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Indeterminate
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| Ettingshausenia
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E. gruenbachiana, E. cf. laevis, Indeterminate
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Platanaceae
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| Extremipollis
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Indeterminate
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Pollen
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| Grebenkia
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G. europeica
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| cf. Grevilleophyllum
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G. constans
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| Hungaropollis
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Indeterminate
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Pollen
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| Juglandiphyllites
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J. pelagicus
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possible affinity with Juglandaceae
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| Krutzschipollites
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Indeterminate
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Pollen
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| Laudaypollis
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Indeterminate
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Pollen
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| Leguminosites
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L. mucronata
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| Longanulipollis
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Indeterminate
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Pollen
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| Menispermites
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M. ettingshausenii, M. summesbergerii
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| Momipetes
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Indeterminate
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Pollen, similar to extant Engelhardtia
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| Myricophyllum
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M. cf. zenkeri, M. serratum
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Affinities with Myrica
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| Myrtacaedites
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Indeterminate
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Pollen, Representative of Myrtaceae
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| Normapolles
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Indeterminate
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Pollen, very abundant. possibly associated with Juglandaceae
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| Oculipollis
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O. sp, O. parvoculus O.zaklinskaiaiae
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Pollen
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| cf. Pandemophyllum
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Indeterminate, P. cf. proteoides
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| Pseudoplicapollis
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Indeterminate
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Pollen
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| Quereuxia
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Q. angulata
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Possible affinity with Trapa
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| Rodgersia
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Indeterminate
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| Semioculipollis
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Indeterminate
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Pollen
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| Suemegipollis
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Indeterminate
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Pollen
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| Ternstroemites
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T. (?) neueweltensis
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| Tricolpopollenites
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Indeterminate
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Pollen, At least 8 distinct taxa
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| Viburniphyllum
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V. ermannii, V. austriacum
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References
- ^ Draganits, Erich; Wagreich, Michael; Hofer, Gerald; Hofmann, Christa-Charlotte; Reischenbacher, Doris; Grundtner, Marie-Louise; Neuhuber, Stephanie; Bottig, Magda (2011). "Stratigraphy and geochemical characterisation of Upper Cretaceous non-marine - marine cycles (Grünbach Formation, Gosau Group, Austria". Austrian Journal of Earth Sciences. 104 (2): 90–107.
- ^ Grundtner, Marie-Louise (2011). Sedimentologie und Sedimentpetrographie der Gosau-Gruppe bei Dreistetten-Muthmannsdorf (Niederösterreichische Kalkalpen) (Master's thesis). University of Vienna. pp. 21–35, 51–57, 71. doi:10.25365/thesis.17461.
- ^ Sachs, Sven; Hornung, Jahn J. (May 2006). "Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper Cretaceous (Lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria)". Geobios. 39 (3): 415–425. doi:10.1016/j.geobios.2005.01.003. ISSN 0016-6995.
- ^ Bunzel, Emanuel (1871). "Die Reptilfauna der Gosau-Formation in der Neuen Welt bei Wiener-Neustadt". Abhandlungen der Kaiserlich-königlichen Geologischen Reichsanstalt. 5: 1–18.
- ^ a b c Seeley, H. G. (1881-02-01). "The Reptile Fauna of the Gosau Formation preserved in the Geological Museum of the University of Vienna: With a Note on the Geological Horizon of the Fossils at Neue Welt, west of Wiener Neustadt, by Edw. Suess, Ph.D., F.M.G.S., &c., Professor of Geology in the University of Vienna, &c". Quarterly Journal of the Geological Society. 37 (1–4): 620–706. doi:10.1144/GSL.JGS.1881.037.01-04.49. ISSN 0370-291X. S2CID 219235284.
- ^ Buffetaut, Eric (1979). "Revision der Crocodylia (Reptilia) aus den Gosau-Schichten (Ober-Kreide) von Österreich". Beiträge zur Paläontologie von Österreich. 6: 89–105.
- ^ Buffetaut, Eric (1989). "Erster nachweis von Choristodera (Reptilia, Diapsida) in der Oberkreide Europas: Champsosaurierwirbel aus den Gosau-Schichten (Campan) Niederösterreichs". Sitzungsberichten der Österreichs Akademis der Wissenschaften Mathematisch-Naturwissenschaftlichen Klasse, Abteilung. 197: 389–394.
- ^ Rabi, Márton; Vremir, Mátyás; Tong, Haiyan (2012-09-01), "Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania)", Vertebrate Paleobiology and Paleoanthropology, Springer Netherlands, pp. 307–336, doi:10.1007/978-94-007-4309-0_19, ISBN 9789400743083
- ^ Buffetaut, Eric; Ősi, Attila; Prondvai, Edina (March 2011). "The pterosaurian remains from the Grünbach Formation (Campanian, Gosau Group) of Austria: a reappraisal of 'Ornithocheirus buenzeli'". Geological Magazine. 148 (2): 334–339. Bibcode:2011GeoM..148..334B. doi:10.1017/S0016756810000981. ISSN 1469-5081. S2CID 131376979.
- ^ Csiki-Sava, Zoltan; Buffetaut, Eric; Ősi, Attila; Pereda-Suberbiola, Xabier; Brusatte, Stephen L. (2015-01-08). "Island life in the Cretaceous - faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago". ZooKeys (469): 1–161. doi:10.3897/zookeys.469.8439. ISSN 1313-2970. PMC 4296572. PMID 25610343.
- ^ Herman, Alexei B.; KVAČEK, JIŘÍ (2007). "Early Campanian Grünbach flora of Austria: systematic composition and palaeoclimatic interpretations". Acta Palaeobotanica. 47 (1): 37–55.
- ^ KVACEK, JIRÍ; HERMAN, ALEXEI (2004). "The Campanian Grünbach Flora of Lower Austria: palaeoecological interpretations". Annalen des Naturhistorischen Museums in Wien. 106 A.
- ^ B., Herman, Alexei (2010). Late Cretaceous Grünbach Flora of Austria. Kvaček, Jiří. Wien: Naturhistorisches Museum. ISBN 9783902421432. OCLC 517260675.
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